The most notable morphological feature of vertebrate teeth is the variability in the crowns of the molars and premolars that appear in different and complex shapes. The basic units of the crowns are cusps or conical projections on the crown of molars and premolars, the number morphology and orientation of the cusps vary greatly, even in the same animal, depending on the feeding habits. These cusped teeth originated from single cusped teeth of ancient mammal like reptiles. The cusps grow on the crown of molars and premolars and are responsible for the great variability there is in teeth even on the same animal and species of vertebrates. The cusps grow the top of the teeth down and have dentine and enamel in them. The evolution of the molar and premolars in vertebrate teeth in mammals follows a system of complexity where in the Pertmain period the ancient reptile mammals had only single cusped teeth but evolved later to cause the development of multi cusped teeth having many patterns
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